Atwaterfactorscaloriesfromproteincarbohydratecaloriesfoodlabelroundingmacronutrientcaloriesnutritionlabelcalculationproteincaloriespergramproteinmetabolismthermiceffectoffood 5 Jul 2026 How Many Calories Are in a Gram of Protein? A gram of protein provides approximately 4 kilocalories of metabolizable energy. Therefore, when …